Updated data on Litopterna from the Huayquerian Stage/Age (Late Miocene-Early Pliocene) of central-east Argentina

Hitherto, only the Proterotheriidae Ameghino, 1887 litopterns from the Cerro Azul Formation (Chasicoan-Huayquerian Stages/Ages; Late Miocene-Early Pliocene) in La Pampa Province had been studied, recognizing Proterotheriidae indet. at Cerro La Bota, Diplasiotherium pampa Soria, 2001 at several localities, Eoauchenia primitiva Ameghino, 1887 at El Guanaco, and cf. Brachytherium cuspidatum Ameghino, 1883 at Salinas Grandes de Hidalgo, which added to Epecuenia thoatherioides Cabrera, 1939 from Laguna Epecuen in Buenos Aires Province; this locality also yielded the Macraucheniidae Gervais, 1855 Huayqueriana cristata (Rovereto, 1914). Now, we enlarge the study to both litoptern families and to the outcrops of the Cerro Azul Formation in Buenos Aires Province. Concerning Macraucheniidae, we recognize: Scalabrinitherium bravardi Ameghino, 1883 at Salinas Grandes de Hidalgo, Telen, and Guatrache (La Pampa), and Laguna Epecuen (Buenos Aires); Paranauchenia denticulata Ameghino, 1891, Promacrauchenia sp., and Cullinia sp. at Salinas Grandes de Hidalgo and Laguna Chillhue (La Pampa); cf. Oxyodontherium zeballosi Ameghino, 1883b at Laguna del Monte (Buenos Aires); and Promacrauchenia sp. at Cantera Relleno Sanitario (Buenos Aires). Referring to Proterotheriidae, new results reveal the first record of Diplasiotherium pampa in Buenos Aires (Arroyo El Venado) and Neobrachytherium sp. at Laguna Epecuen and Laguna La Paraguaya (Buenos Aires). The litoptern diversity is close to that from the Lower Member of the Ituzaingo Formation, Late Miocene, Entre Rios Province. The litoptern record does not provide confident data to support or reject the established relative biostratigraphic succession of localities of the Cerro Azul Formation, which is mainly based on some rodent lineages.