Genetic coupling of life-history and aerobic performance in juvenile 1 Atlantic salmon

3 4 Short title: Genetic coupling of life-history and metabolism 5 6 7 Jenni M. Prokkola1*, Eirik R. Åsheim1, Sergey Morozov1, Paul Bangura1, Jaakko Erkinaro2, 8 Annukka Ruokolainen1, Craig R. Primmer1,3, Tutku Aykanat1 9 10 1Organismal and Evolutionary Biology Research Programme, University of Helsinki, Finland 11 2Natural Resources Institute Finland 12 3Institute of Biotechnology, Helsinki Institute of Life Science (HiLIFE), University of 13 Helsinki, Finland 14 *Corresponding author, PO Box 65, FI-00014 University of Helsinki, Finland, 15 jenni.prokkola@helsinki.fi 16 17 Acknowledgements 18 We thank Nikolai Piavchenko, Katja Maamela, Markus Lauha, Petra Liljeström, Suvi Ikonen, 19 Mikko Immonen, and Anna Toikkanen for fish care and technical help during the experiment, 20 and Lammi Biological Station for a high-quality research environment. Dr. Markus Haapala 21 and the Nordic University Hub NordicPOP (Nordforsk, project no. 85352) are acknowledged 22 for the 3D printing service, and Heidrikur Bergsson for sharing the chamber cap model. The 23 study was funded by Academy of Finland (T. Aykanat: 325964, 1328860, C. R. Primmer: 24 284941, 286334, 314254 and 314255), the European Research Council under the European 25 Articles Union’s Horizon 2020 research and innovation program (grant no. 742312), and the 26 University of Helsinki. 27