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The Rockefeller University Press $30.00 J. Cell Biol. Vol. 207 No. 2 299–315 www.jcb.org/cgi/doi/10.1083/jcb.201404067 JCB 299 Correspondence to James E. Bear: jbear@email.unc.edu Abbreviations used in this paper: 4-OHT, 4-hydroxy-tamoxifen; AMPK, AMPactivated protein kinase; ANOVA, analysis of variance; BRSK, brain-specific kinase; DN, dominant negative; FMI, forward migration index; MARK, microtubule affinity-regulating kinase; MMP, matrix metalloproteinase; NUAK, nua kinase; PDMS, polymethylsiloxane; qRT-PCR, quantitative RT-PCR; SIK, salt-inducible kinase; YAP, Yes-associated protein. Introduction