Simon Hugh Piper Maddrell, Scd, Frs, 1937-2020 Obituary

Helen Skaer, Michael J O'Donnell,Julian A T Dow

JOURNAL OF EXPERIMENTAL BIOLOGY（2020）

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![Graphic][1] \n\nSimon Hugh Piper Maddrell – Hugh after his father and Piper, of which he was very proud, after the sandpipers his parents heard on their camping honeymoon – was born in 1937, the oldest of five siblings. His upbringing on his parents\u0027 farm meant that nature ran deep in his

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