The Aquilegia genome provides 1 insight into adaptive radiation and 2 reveals an extraordinarily 3 polymorphic chromosome with a 4 unique history 5

*For correspondence: scott.hodges@lifesci.ucsb.edu (SH); magnus.nordborg@gmi.oeaw.ac.at (MN) 1Gregor Mendel Institute, Austrian Academy of Sciences, Vienna BioCenter (VBC), Vienna, 11 Austria; 2Department of Ecology, Evolution, and Marine Biology, University of California 12 Santa Barbara, Santa Barbara, California, USA; 3Central-European Institute of Technology 13 (CEITEC), Masaryk University, Brno, Czech Republic; 4Vienna Graduate School of 14 Population Genetics, Vienna, Austria; 5Department of Energy Joint Genome Institute, 15 Walnut Creek, California, USA; 6HudsonAlpha Institute of Biotechnology, Huntsville, 16 Alabama, USA; 7Institute of Experimental Botany, Centre of the Region Haná for 17 Biotechnological and Agricultural Research, Olomouc, Czech Republic; 8Department of 18 Organismic and Evolutionary Biology, Harvard University, Cambridge, Massachusetts, USA 19