1 Voluntary exercise improves metabolic profile in high-fat fed glucocorticoid treated rats 1 2

1 School of Kinesiology and Health Science, Faculty of Health, Muscle Health Research Center and 5 Physical Activity and Chronic Disease Unit, York University, 4700 Keele St., Toronto, ON, Canada, M3J 6 1P3 7 *Co-AuthorsBoth identified authors contributed equally to this entire manuscript. 8 9 **Corresponding Author: Dr. Michael Riddell, School of Kinesiology and Health Science, Faculty of 10 Health, Muscle Health Research Center and Physical Activity and Chronic Disease Unit, York University, 11 4700 Keele St., Toronto, ON, Canada. Tel: 416-736-2100 Fax: 416-736-5774. Email: mriddell@yorku.ca 12