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His best‐known contributions in about 250 research papers and 14 books deal with environmental plant physiology and ecology, especially in desert plants. His recent research dealing with the interaction of light with high temperature, water and nutritional stresses, and the underlying physiological and biochemical mechanisms responses to excess light, has been supported by a Forschungspreis from the Alexander von Humboldt Foundation and by grants from the US DOE. In retirement he is continuing research into photosynthetic processes in old leaves within the deeply shaded canopies of woody plants and development of new noninvasive remote sensing instruments.
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